Operating at sea level to 3000 ft means the air is more dense, so the engine will make more power, which requires less throttle/fuel consumption to move the rig down the road.
If one looks solely at how gear ratio affects fuel economy, the amount of applied throttle (fuel consumed) to get the rig moving down the road/maintain cruising speed is more impactful to MPG than cruising RPM.
I've run 4:56's with 33's/5 speed and 4:56's /AW4 and my experience is that 4:56's/33's are great with a 5speed, but not low enough gearing with an AW4.
On a prior XJ, I was running 4:56's/5speed/33's, but swapped out the BA10/5 manual for an AW4 auto and after the trans swap, my rig felt like my axle gear ratio was back to 4:10's.
The trans would constantly shift out of OD into 3rd on the slightest grade, would hunt between 3rd an OD when cruising and I was required to apply a lot more throttle to get the rig up to speed and maintain cruising speed, which resulted in a 3-5 MPG loss in fuel economy.
I've experienced none of these issues running 4:88 gearing, which has been awesome with an AW4 and 33's.